Evolutionary Significance of Pylentonemid Radiolarians and Their Late Devonian Species from Southwestern Tianshan,China  

作　　者：LIU Yu HAO Shougang 

机构地区：[1]Department of Geology, Peking University, Beijing 100871

出　　处：《Acta Geologica Sinica(English Edition)》2006年第5期647-655,共9页地质学报（英文版）

基　　金：This work was supported by the National Nature Science Foundation of China(No.40232019).

摘　　要：A group of radiolarian fossils with a complete morphological gradient of increasing polarization are reported from upper Devonian successions in southwestern Tianshan, China. Specifically, the important transitional mophotypes, Pylentonema transitorum sp. nov., Archocyrtium medium sp. nov. and Quadrapesus transitivus sp. nov, were found. Based on the fact that the new transitional radiolarian fossils resemble more closely to pylentonemids （cyrtoid nassellarians） than to entactinids （spumellarians） in morphology, they are assigned to pylentonemids. Pylentonemids share characters of both the Nassellaria （e.g., horned cephalis, basal opening with pylome） and Spumellaria （e.g., system of trabecular spicules）, and they are thus of evolutionary significance. The evolutionary trend from spumellarians to nassellarians is characterized by （1） internal spicule： from simple trabecular spicules to complicate components （A;Ir,II;D;V;Lr,Li;MB）; （2） polarization of the radiolarian tests due to rearrangment of spine, from radial to an axial symmetry; （3） the gradual formation of a basal opening （aperture, pylome, or podome）; and the transition from a pseudopylome, a primative pylome with a narrow margin on the outer shell, to a true pylome rimmed with a narrow, elevated impermite wall.A group of radiolarian fossils with a complete morphological gradient of increasing polarization are reported from upper Devonian successions in southwestern Tianshan, China. Specifically, the important transitional mophotypes, Pylentonema transitorum sp. nov., Archocyrtium medium sp. nov. and Quadrapesus transitivus sp. nov, were found. Based on the fact that the new transitional radiolarian fossils resemble more closely to pylentonemids （cyrtoid nassellarians） than to entactinids （spumellarians） in morphology, they are assigned to pylentonemids. Pylentonemids share characters of both the Nassellaria （e.g., horned cephalis, basal opening with pylome） and Spumellaria （e.g., system of trabecular spicules）, and they are thus of evolutionary significance. The evolutionary trend from spumellarians to nassellarians is characterized by （1） internal spicule： from simple trabecular spicules to complicate components （A;Ir,II;D;V;Lr,Li;MB）; （2） polarization of the radiolarian tests due to rearrangment of spine, from radial to an axial symmetry; （3） the gradual formation of a basal opening （aperture, pylome, or podome）; and the transition from a pseudopylome, a primative pylome with a narrow margin on the outer shell, to a true pylome rimmed with a narrow, elevated impermite wall.

关 键 词：RADIOLARIAN spumellarian nassellarian entactinids pylentonemids phylogenetic evolution Late Devonian southwestern Tianshan 

分 类 号：Q915.2[天文地球—古生物学与地层学]