杉木次生韧皮部筛胞发育中的超微结构变化  

Ultrastructure change during the development sieve cell of secondary phloem in Cunninghamia lanceolata

作　　者：彭冶[1] 樊汝汶[1] 黄金生[1] 甘习华[1] 方炎明[1] 

机构地区：[1]南京林业大学森林资源与环境学院,江苏南京210037

出　　处：《南京林业大学学报（自然科学版）》2008年第6期47-51,共5页Journal of Nanjing Forestry University：Natural Sciences Edition

基　　金：国家自然科学基金资助项目(39730350)

摘　　要：杉木筛胞发育分化过程中，超徽结构表现出以下特征：发育分化初期，筛母细胞与形成层细胞极为相似，细胞壁薄，仅有胞间层和累积少量壁物质的初生壁；随后伴随筛咆的径向扩张，初生壁开始加厚，壁密度、厚度均一，没有明显的分层现象，未表现出异质性，仍为初生壁；最终，径向直径约为形成层细胞的1～1．5倍。伴随筛母细胞的细胞壁增厚，前质体多分化为淀粉质雄，P-质雄较少；筛域的分化与筛胞壁的发育同步；内质网、高尔基体参与筛域的形成；成熟筛胞的筛域多见于其端部，侧壁较少。由此可见，杉科在裸子植物中属于演化水平较高的类群；参与筛胞分化的细胞器主要是内质网，其次是线粒体和高尔基体；筛胞分化中，细胞核的变化特征与细胞程序性死亡（PCD）极为相似。The process of developing the sieve cell of secondary phloem in Cunninghamia lanceolata （ Lamb. ） Hook. could be divided into two stages, i.e. radial expanding of the mother cell and orderly degradation of nuclei and organelle. After first stage, the sieve area began to form, and the wall became tiny thicken, but sill was primary cell wall. In the second stage, organelles degraded. The endoplasmic reticulum played a main role in the whole process. The vacuole membrane was not ruptured. The mature sieve cell plastid was starch. The P - protein was not found. The nuclear degradation was similar to program cell death.

关 键 词：杉木 筛胞 发育 超微结构 

分 类 号：S718[农业科学—林学]