A New Family of Moths from the Middle Jurassic(Insecta:Lepidoptera)  

作　　者：HUANG Diying Andre NEL Joeel MINET 

机构地区：[1]Nanjing Institute of Geology and Palaeontology, State Key Laboratory of Palaeobiology and Stratigraphy, Chinese Academy of Sciences, Nanjing, 210008, China [2]UMR CNRS 7205, Museum National d＇Histoire Naturelle, Departement Systematique et Evolution, C.P. 50, Entomologie, F-75231 Paris Cedex 05, France

出　　处：《Acta Geologica Sinica(English Edition)》2010年第4期874-885,共12页地质学报（英文版）

基　　金：the National Natural Science Foundation of China(grants nos.40672013 and 40632010);the Chinese Academy of Sciences(grant no.KZCX2-YW-QN104);the Major Basic Research Projects of China's Ministry of Science and Technology(no.2006CB806400)

摘　　要：Three lepidopteran species, from the Middle Jurassic Daohugou beds （inner Mongolia, China）, are described in a new family, Mesokristenseniidae, and new genus, Mesokristensenia, which could represent the sister group of the Micropterigidae. Mesokristensenia differs from all extant Lepidoptera, but one genus （Agathiphaga, Agathiphagidae）, in retaining four median veins in the forewing, a plesiomorphy also present in many Trichoptera. Evidence for placing Mesokristensenia in the Lepidoptera includes four traits, notably a previously unrecorded autapomorphy of this insect order： beyond stem Ml＋2, vein M1 is bent and connected to cross-vein r-m （in both wing pairs）. Among 24 characters taken into account to assess the systematic position of Mesokristensenia, 12 are considered informative for a cladistic analysis involving this fossil taxon and the four suborders recognized in present-day Lepidoptera （Zeugloptera, Aglossata, Heterobathmiina, and Glossata）.Three lepidopteran species, from the Middle Jurassic Daohugou beds （inner Mongolia, China）, are described in a new family, Mesokristenseniidae, and new genus, Mesokristensenia, which could represent the sister group of the Micropterigidae. Mesokristensenia differs from all extant Lepidoptera, but one genus （Agathiphaga, Agathiphagidae）, in retaining four median veins in the forewing, a plesiomorphy also present in many Trichoptera. Evidence for placing Mesokristensenia in the Lepidoptera includes four traits, notably a previously unrecorded autapomorphy of this insect order： beyond stem Ml＋2, vein M1 is bent and connected to cross-vein r-m （in both wing pairs）. Among 24 characters taken into account to assess the systematic position of Mesokristensenia, 12 are considered informative for a cladistic analysis involving this fossil taxon and the four suborders recognized in present-day Lepidoptera （Zeugloptera, Aglossata, Heterobathmiina, and Glossata）.

关 键 词：TRICHOPTERA LEPIDOPTERA Mesokristenseniidae wing venation PHYLOGENY China Middle Jurassic 

分 类 号：Q969.42[生物学—昆虫学] Q915.819.7[天文地球—古生物学与地层学]