We thank S.-Y. Wang, H.-B. Liang, L.-J. Zhang and T.-H. Luo (Institute of Zoology, Chinese Academy of Sciences) for some species identification; John P. Ball (Swedish University of Agricultural Sciences, Umea, Sweden), Joshua M. Jacobs (Department of Renewable Resources, University of Alberta) and anonymous referees for con- structive suggestions to improve the manuscript; and the staff of Fengtongzai Natural Reserve and Wolong Natural Reserve, Sichuan province, for assistance in field investigations. This study was supported by National Natural Science Foundation of China (NSFC-30570215, 30670285), KFBG (Kadoorie Farm & Botanic Garden) Biodiversity Studentship of Hongkong, International Foundation for Sciences (D/3684-1, IFS, Stockholm, Sweden), and National Science Fund for Fostering Talents in Basic Research (Special subjects in animal taxonomy, NSFC- J0630964/J0109).
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