the National Natural Science Foundation of China(31900336);the National Basic Research Program of China(2007CB109106);the Key Research Programs in Colleges and Universities of Henan Province(16A180039 and 19A180032);the China Postdoctoral Science Foundation(2016M592304).
Spatial patterns in heterogeneity are generally deemed a central causal factor influencing the physiology and behavior of animals in ecological systems.However,knowledge remains limited about how such patterns influen...