supported by the National Basic Research Program of China(Grant No.2006CB806400);the Chinese Academy of Sciences(Grant No.KZCX2-EW-106);the Basic Work Program of MST of China(Grant No.2006FY120300-15);the National Natural Science Foundation of China(Grant No.40532010);Special Fund for Fossil Excavation and Preparation of Chinese Academy of Sciences;US National Science Foundation(Grant No.EAR-0120727,BCS-0309800,BCS-0820602)~~