supported by the National Natural Science Foundation of China (41172018, 41572020, 41688103);the Major Research Plan of the National Natural Science Foundation of China (91514302);the Strategic Priority Research Program (B) of the Chinese Academy of Sciences (XDB18030504);the National Key Basic Research Program of China (2012CB821900);the National Science Fund for Distinguished Young Scholars (40825005);the Laiyang Government Cooperation Dinosaur Project~~