小草蔻花部维管束系统的解剖学研究  

Studies on Vascular System Anatomy of the Flower of Alpinia henryi

作　　者：唐源江[1] 谢中誉[1] 廖景平[1] 

机构地区：[1]中国科学院华南植物研究所,广东广州510650

出　　处：《热带亚热带植物学报》2002年第1期27-32,共6页Journal of Tropical and Subtropical Botany

基　　金：国家自然科学基金项目(39870087)资助

摘　　要：小草蔻花梗的维管束可分成外环和中心区两部分。进入室下区后,外环维管束向外偏移,其数目由约13束增加至约40束;中心区由不规则分布的18束变为三角形排列的12束。延伸至子房室区时,外环基本不变,而原来中心区的维管束则成为3束心皮背束、3束隔膜束及5-7束小的胎座维管束。3束心皮背束外方的主支经延长部最后进入花萼,胎座维管束于延长部基部消失。及至延长部基部,3束隔膜束的内侧横向分支,并最后与部分外环的维管束及心皮背束内方的细小分支在此形成一个维管束网结,而外侧的主支直接向上延伸;近轴面的隔膜束主支分裂成4支最后进入仅有的1枚功能雄蕊,远轴面的2束隔膜束主支则各分成2-3束最后进入唇瓣。延长部的维管束网结部分延伸入花瓣、唇瓣和腺体基部,少量进入花柱。首次提出姜科植物的花萼既代表了3枚花萼片,又包含了3枚缺失的外轮雄蕊;支持姜科的唇瓣代表了两枚缺失的内轮雄蕊及两枚腺体是隔膜腺变异结构的观点。Outerring(OR)andcentralregion(CR)of vascularbundlesin pedicelof Alpinia henryi can berecognized.WhenORandCRextendintosublocularregion,thebundlesof ORmovestowardoutside andincreasesinnumberfrom13to40,scatteredCRbundlesarearrangedintotriangleandtheirnumber variesfrom18into12.Atthelocularregion,theCRbundlesarearrangedas3carpellarydorsalbundles,3parietalbundles,and5-7smallplacentalbundleswhichfadeawayat thebaseof prolongation.Outer largebranchesof thethreecarpellarydorsalbundlesfinallyenterthecalyx.Atthebaseof theprolongation,threeparietalbundlesproducesometransversebrancheswhichbecomevascularplexuswithbranchesof partialbundlesand a few smallbranchesoriginatedfromthreecarpellarybundles,buttheirbundles extenddirectlyupwards.Finally,theadaxialparietalbundlesplitsup into4brancheswhichrunsup into a functionalstamen,whilethetwoabaxialparietalbundlessplitup respectivelyinto2-3branchesgoing to thelabellum.Thebundlesof thevascularplexusfinallyenterpetals,labellumandstyle.It suggested thatthecalyxcontainsnot onlybundlesin threesepalsbutalsoin threeabsentstamens.The authors supporttheviewsthatthelabellumrepresentstwo absentstamensof innerwhorlof androeciumandthe twoglandsareof modifiedgynopluralnectary.

关 键 词：小草蔻 花 维管束系统 解剖学 姜科 

分 类 号：Q944.58[生物学—植物学] Q949.718.3