Acknowledgements This study was supported by the National Natural Science Foundation of China (Grant No. 31070182). The authors of this paper are grateful to the administrators of the herbaria PE, PYU, K, KATH, and TNS for checking specimens. The authors also thank Professor Dr. H. SCHNEIDER for critical reading of the manuscript and offering helpful suggestions for improving the manuscript, and thank Mr. N. SHRESTHA and Mr. J. W. LI for improving the English.
The species delimitation of Lemmaphyllum, including the former segregated Lepidogrammitis in China, was considered as unresolved. Previous treatments accepted between one and 20 species. In the present study, multivar...