the Knowledge Innovation Program of the Chinese Academy of Sciences(KZCX2-EW-106);the Major Basic Research Projects of MST of China(2012CB821900);the National Natural Science Foundation of China(40532010,40802009);the U.S. National Science Foundation(EAR-0120727,BCS-0309800);中国科学院化石发掘与修理特别支持费资助~~