supported by the National Natural Sciences Foundation of China (Grant 30830017,30970391);National Science Funds for Distinguished Young Scientists (31025024);National Science Fund for Fostering Talents in Basic Research (J0930004);a grant (O529YX5105)from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences
funded by the National Natural Sciences Foundation of China(30830017 and 30970391);CAS Innovation Program(KSCX2-YW-Z-011)to QIAO,Ge-Xia;National Science Fund for Fostering Talents in Basic Research(Special subjects in animal taxonomy,NSFC-J0630964/J0109);a grant(O529YX5105)from the Key Laboratory of the Zoological Systematics and Evolution of the Chinese Academy of Sciences