supported by the National Natural Science Foundation of China(NSFC-32200369);the Science and Technology Project Foundation of Guizhou Province([2020]1Z014);the Key Laboratory Project of Guizhou Province([2020]2003);the Open Project of Ministry of Education Key Laboratory for Ecology of Tropical Islands,Hainan Normal University,China([HNSF-OP-202201]);the Doctoral Research Foundation of Tongren University(trxyDH2102).