partly supported by the Natural Science Foundation of Liaoning Province(20170540642);the General Project of Scientific Research of the Education Department of Liaoning Province(L2015359);the Key Project of the Qinghai Qilian Mountain Nature Reserve Administration(QHTX-2021-006)。
General Project of Scientific Research of Educational Commission of Liaoning Province Under Grant(L2015359);Mentoring Program of Natural Science Foundation of Liaoning Province Under Grant(20170540642)