supported by the National Natural Science Foundation of China (41006084,31172053,31061160187);Natural Science Foundation of Guangdong Province of China (10152408801000013)
supported by the Special Foundation of Educational Commission of Guangdong Province for Talent Introduction(0909128);Projects of International(Region)Cooperation and Exchanges NSFC(31061160187);Special Fund for Agro-scientific Research in the Public Interest(201003068)