the financial support from the National Natural Science Foundation of China(Nos.52172067,92160202);Natural Science Foundation of Guangdong Province,China(Nos.2021B1515020038,2020B1515020036);Guangdong Special Support Program,China(No.2019BT02C629);Guangdong Academy of Sciences Program,China(No.2020GDASYL20200104030)。