Suppated by the National Natural Science Fundation of China(20704042,60225004);the Shanghai City Committee of Science and Technology(06XD14020,07 JC14058,0752nm016);Shanghai Pujiang Talent Plan(07PJ14095);the Know ledge Innovation Program of the Chinese A-cademy of Sciences(KSCX2-YW-G-042);National Basic Research Program of China(2010CB933700)