This work was funded by grants from the National Natural Science Foundation of China (NSFC) (Grant No. 31210103916 and 91019019), Chinese Ministry of Science and Technology (Grant No. 2011CB504206) and Chinese Academy of Sciences (CAS) (Grant Nos. KSCX2-EW-R-02 and KSCX2-EW-J-15) and stem cell leading project XDA01010303 to J.D.J.H.H.N. was supported by the Chinese Academy of Sciences Fellow- ship for Young International Scientist [Grant No. 2012Y1SB0006] and the National Natural Science Foundation of China [Grant No. 31250110524]. The authors thank Dr. Jerome Boyd-Kirkup for extensive editing and Hamna Anwar for proofreading the manu- script.
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