Key Program of the National Natural Science Fund(31130061);National Nature Science Foundation of China(31572278; 31801981;31470214);Central Finance Forestry Protection Grant Funds(2014-2016);Foundation of Shaanxi Academy of Science,China(2016K-20;2018nk-18);the Strategic Priority Research Program of the Chinese Academy of Sciences(XDB31020302);Key Research and Development Program of Shaanxi Province of China(2018PT-04; 2018 ZDXM-NY-049)