supported by Hainan Province Science and Technology Special Fund(ZDYF2021GXJS207,ZDYF2020037,and 2020207);the National Natural Science Foundation of China(22109034,22109035,52164028,and 62105083);the Foundation of State Key Laboratory of Marine Resource Utilization in South China Sea(Hainan University,MRUKF2021029);the specific research fund of the Innovation Platform for Academicians of Hainan Province;the“Young Talent Support Plan”of Xi’an Jiaotong University。