supported by Key Lab of Freshwater Biodiversity Conservation,Ministry of Agriculture(LFBC0807);the Educational Commission of Jiangxi Province of China(GJJ11718);the Natural Science Foundation of Jiangxi Province of China(20144BAB2150004);the Technology Bureau of Ji’an City of China([2012]32-8);the Doctoral Starting up Foundation of Jinggangshan University;China(jz10040)