supported by the Yunnan Digitalization,Development and Application of Biotic Resource(202002AA100007);the National Natural Science Foundation of China(82222064,81973147,82001986,and 82073569);Shandong Medical and Health Technology Development Project(202005010068);Shandong University Distinguished Young Scholars;the Outstanding Youth Science Foundation of Yunnan Basic Research Project(202001AW070021,and 202101AW070001);the Reserve Talent Project for Young and Middle-Aged Academic and Technical Leaders(2012005AC160023);the Key Science Foundation of Yunnan Basic Research(202101AS070040);the Innovative Research Team of Yunnan Province(202005AE160002);the Science Foundation of Yunnan Basic Research(202201AT070010)。