supported by the Guizhou Provincial Key Technology R&D Program (Qian Ke He Zhi Cheng [2021] Generally 200);the Youth Science and Technology Talent Development Project from Guizhou Provincial Department of Education (QJHKYZ[2022]345);the National Natural Science Foundation of China (31960716);the Post-grant of the National Natural Science Foundation of China;Guizhou Academy of Agricultural Science ([2021] 49)。