国家现代农业产业技术体系建设专项(CARS-20);国家公益性行业(农业)科研专项(201503119);福建省高等学校新世纪优秀人才支持计划项目(JA14095)资助;supported by the China Agriculture Research System(CARS-20);the Special Fund for Agro-Scientific Research in the Public Interest(201503119);the Program for New Century Excellent Talents in Fujian Province University(JA14095)