National Natural Science Foundation of China(21276035);Sub-project of Central Sharing Funds for Using Sea Area(2013-348-7);Special Foundation for Ocean Environmental Protection of Ocean and Fisheries Department of Liaoning Province(2012-Inhyhbc-0004,2012-Inhyhbc-0005);Scientific Research Project of Education Department of Liaoning Province(L2013203);Natural Science Foundation of Liaoning Province(2014025014)