This research was supported by the National Key Research and Development Program of China(2022YFC3201702);National Natural Science Foundationof China(U2240226,42201146);Sanjiangyuan National Park Joint Research Program of Chinese Academy of Sciences and The People's Government of Qinghai Province(LHZX-2020-10-3);Science and Technology Project of Sichuan Province(2022NSFSC1001).