supported by the Strategic Priority Research Program of the Chinese Academy of Sciences [grant number XDA20060500];the National Natural Science Foundation of China[grant numbers 41731173 and 42275035];the Natural Science Foundation of Guangdong Province;China [grant number 2022A1515011967];the Science and Technology Program of Guangzhou;China [grant number 202002030492];the Open Fund Project of the Key Laboratory of Marine Environmental Information Technology,the Key Laboratory of Marine Science and Numerical Modeling,Ministry of Natural Resources of the People’s Republic of China [grant number 2020-YB-05];the MEL Visiting Fellowship [grant number MELRS2102];the Independent Research Project Program of the State Key Laboratory of Tropical Oceanography [grant number LTOZZ2005];the Key Special Project for the Introducing Talents Team of the Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou)[grant number GML2019ZD0306];the Innovation Academy of South China Sea Ecology and Environmental Engineering [grant number ISEE2018PY06]