Strategic Priority Research Program of the Chinese Academy of Sciences(XDA13020206);Innovation Academy of South China Sea Ecology and Environmental,Engineering,Chinese Academy of Sciences(ISEE2018PY03,ISEE2018ZD02);Science and Technology Service Network Initiative of Chinese Academy of Sciences(KFJ-STS-ZDTP-055);Science and Technology Planning Project of Guangdong Province,China(2017B030314052)。