Supported by the National Natural Science Foundation of China(31500013,30770013);the Biodiversity Survey and Assessment Project of the Ministry of Ecology and Environment of China(2019HJ2096001006);the Provincial Key Technologies R&D Program of Guizhou[(2017)2513];the Earmarked Fund for Hebei Edible Fungi Innovation Team of Modern Agro-industry Technology Research System(HBCT2018050205);Special Foundation for Key Agricultural Generic Technology Development(16227303D).