National Natural Science Foundation of China(41976112,31971432,41576125);Fund for strategic biological resources,Chinese Academy of Sciences(KFJ-BRP-017-48);Key Special Project for Introduced Talents Team of Southern Marine Science and Engineering Guangdong Laboratory(Guangzhou)(GML2019ZD0401,GML2019ZD0405);Science and Technology Basic Resources Investigation Program of China(2017FY201404)。