financially supported by the Program for New Century Excellent Talents in University(NCET-07-0445);the Ministry of Education of China,and the major project of the Natural Science Foundation of the Jiangsu Higher Education Institutions of China(07KJA18016)
sponsored by a grant from the 46th Chinese postdoctoral science foundation,the Natural Science Foundation of China(31101629 and 31172065);the Key Laboratory of Zoological Systematics and Evolution of the Chinese Academy of Sciences(O529YX5105)