Supported by the Ocean Industry Research Special Funds for Public Welfare Projects(201305022-6);the National Science and Technology Support Program Project of China(2012BAD25B02,2013BAD10B01-2-3);the Beijing Earmarked Fund for Modern Agro-industry Technology Research System(SCGWZJ20121104-4);the Special Fund for Basic Research of the Chinese Academy of Agricultural Sciences(2013ZL043);the Central Public-Interest Scientific Institution Basal Research Fund(2013ywf-sl-4)~~