Supported by the National Natural Science Foundation of China (31750001,31670015);the Biodiversity Survey and Assessment Project of the Ministry of Ecology and Environment,China (2019HJ2096001006)。
This study was supported by the Knowledge Innovation Programof the Chinese Academy of Sciences (KSCX2-YW-G-025);the National Natural Science Foundation of China (Nos 30525002,30420120049);The field work was supported by the National Science Foundation of the USA(DEB-0321846)