supported by the National Key Research Development Program of China(2018YFA0800900,2016YFA0500100,and 2018YFA0800700);the National Natural Science Foundation of China(31671402,91954108,91857201,31671233,31701018,and U1702288);the Strategic Priority Research Program of the Chinese Academy of Sciences(XDA12040218)。
With the rapid development of high-throughput sequencing technologies,it has recently become evident that the eukaryotic genome is extensively transcribed[1,2].Large-scale projects for the systematic annotation and fu...