supported by the Funds of National Natural Sciecne Foundation of China(41106143);Guangdong Natural Science Foundation(S2011040000377);China Postdoctoral ScienceFoundation(2011M500133)
supported by the Funds of the Chinese Academy of Sciences for Key Topics in Innovation Engineering(KZCX2-YW-216,KSCX2-YW-G-065,KSCX2-YW-G-073);Natural Science Funds of South China Sea Institute of Oceanology for Young Scholar(SQ200903);Funds of Frontier Areas of SCSIO (LYQY200805);Open Project Program of Key Laboratory of Marine Bio-resources Sustainable Utilization(LMB091013)