国家自然科学基金项目(31271641;31461143015;31471438);江苏省农业三新工程项目(SXGC[2014]313);江苏高校优势学科建设工程资助项目(PAPD);江苏省普通高校研究生科研创新计划项目(KYZZ_0364);扬州大学高端人才支持计划项目(2015-01)资助;supported by the National Natural Science Foundation of China(31461143015;31271641;31471438);Jiangsu "Three-innovation" Agricultural Project(SXG2014313);the Priority Academic Program Development of Jiangsu Higher Education Institutions(PAPD);Innovation Research Program for Graduate Students for Higher Education of Jiangsu Province(KYZZ_0364);and the Top Talent Supporting Program of Yangzhou University(2015-01)