financial support from the Foundation of Guizhou Province, China (Grant No.[2014] 7619);the Teaching Reform and Research Project of the Guizhou University, China (Grant No.JG 2013014); the science and technology innovation foundation of the Guizhou University (Grant No.2016009); the opening foundation of the Key Laboratory of Green Pesticide and Agricultural Bioengineering, Ministry of Education, Guizhou University (Grant No. 2016GDGP0102)~~