Supported by the Outstanding Research Achievement Award Project of Tianjin Agricultural University;the National Natural Science Foundation of China(21875117,31572492);the Innovative and Entrepreneurial Training Plan for Tianjin College Students(202010061066);the Veterinary“Innovative Talent People of Young and Middle Age Key Members Project”of Tianjin;the Natural Science Foundation of Tianjin(18JCYBJC30100)。