supported by the National Key Research and Development Program of China(2022YFD2400101);the National Natural Science Foundation of China(31930111 and 32102785);the Strategic Priority Research Program of Chinese Academy of Sciences(XDA024030104);the China Agriculture Research System of Ministry of Finance and Ministry of Agriculture and Rural Affairs;the Special Research Assistant Project CAS;the China Postdoctoral Science Foundation(2021M693344);the Excellent Young Talents program of Wuhan。