The Joint Funds of the National Science Foundation of China and Yunnan Province Government(No.U1202262);the National Natural Science Foundation of China(No.30470011,31270075);International Cooperation Yunnan Program of Innovation to Strengthen Provinces by Science&Technology(No.2009AC013);Key Laboratory for Plant Diversity and Biogeography of Eastern Asia,Kunming Institute of Botany,Chinese Academy of Sciences(No.0806361121)