supported by National Key R&D Program of China(2021YFF0704500,2021YFF0703701,and 2022YFC3400405);the Strategic Priority Research Program of the Chinese Academy of Sciences(XDB38040300);the 14th Five-year Informatization Plan of Chinese Academy of Sciences(CASWX2021SF-0203);the National Natural Science Foundation of China(91940306,31871294,31970647,81902519,and 32200478);the Special Investigation on Science and Technology Basic Resources of the MOST,China(2019FY100102);the China Postdoctoral Science Foundation(2022M713311);the National Genomics Data Center,China。