This work was supported by the Project of China Scholarship Council(Grant No.LJF[2017]50863);the Program for National Innovation Training of Undergraduate(Grant No.2018109760007,2018109760008);the Program for First-class Discipline Construction in Guizhou Province(Grant No.QJKYF[2017]85);the Project for Science and Technology of Guizhou Province(Grant No.QKHJC[2017]1004);the Project for Major Research by Innovation Group in Guizhou Education Department(Grant No.QJHKY[2018]027);the Project for Provincial Key Discipline of Ecology(Grant No.ZDXK[2013]08);the Project for Cooperative Base for Production,Education and Research of Organic Vegetable Ecological Cultivation and Breeding(Grant No.KY[2013]125);the Program for Innovative Training of Undergraduate in Guizhou(Grant No.2018520777,2018520778,2018520831,2018520861,20195200341,20195201373);the Program for Educational Incubation of Guiyang University(Grant No.GYU-KJT[2019]-06,GYU-KJT[2019]-07);the Discipline and Master's Site Construction Project of Guiyang University by Guiyang City Financial Support Guiyang University(Grant No.SH-2020).