东北农业大学大豆生物学教育部重点实验室开放基金项目(SB14A04);国家现代农业产业技术体系建设专项(CARS-04-CES18);国家自然科学基金项目(31501329);山东省自然科学基金项目(ZR2015YL070)资助;supported by the Open Foundation of Key Laboratory of Soybean Biology of Ministry of Education;Northeast Agricultural University(SB14A04);the China Agriculture Research System(CARS-04-CES18);the National Natural Science Foundation of China(31501329);the Natural Science Foundation of Shandong Province(ZR2015YL070)