financially supported by the National Natural Science Foundation of China(22005149,21975129,and 22172077);the Natural Science Foundation of Jiangsu Province(BK20200777 and BK20211573);the Natural Science Foundation of the Higher Education Institutions of Jiangsu Province,China(20KJB430034);the Science Fund for Distinguished Young Scholars,Nanjing Forestry University(JC2019002)。